Changes in the Physical-Chemical Environment
Upstream Effects
The major upstream effect of the dam is the creation of a reservoir or a controlled stretch of river. In general, a considerable body of water is stored in the reservoir behind the dam. When water is stored for some time in large reservoirs it may become stratified, whereby the warmer waters (that are less dense) float on the cooler (more dense) waters in the deeper parts of the reservoir. The surface waters are further warmed by the sun and, as the density of water decreases with rising temperatures, mixing of the cooler mass of water at the bottom is prevented. As a result, stratification of the water column may occur.

In these circumstances, there is a strong seasonality whereby the water mass at the bottom may become deoxygenated and unable to support life, although the surface waters may be well oxygenated. The lake may or may not return to normal when storms in the monsoon cause the water at the bottom to mix with that at the surface, and produce a more uniform concentration of oxygen. Deep reservoirs may become permanently stratified and de-oxygenated. New reservoirs show an increased tendency to become anoxic in the period immediately after filling when the large quantities of vegetation flooded by the rising water start to decay. This phase may last for several months or even years, particularly where the original area covered by the reservoir was forested. The tendency for reservoirs to become deoxygenated is also accelerated in regions where there is a substantial input of nutrients from agricultural fertilizers, animal wastes or human sewage, either directly into the reservoir or into the river feeding it.

The warm upper waters of a reservoir are well lit by the sun's rays and, in contrast to the swiftly flowing river waters, can support abundant phytoplankton (see Section 6) that produce oxygen by photosynthesis. In contrast, the deeper waters receive little light, and phytoplankton are scarce. Organic matter – mainly FPOM – carried by the river is deposited on the bottom of the reservoir and provides an energy source for microbes – especially bacteria (see Section 6). They consume oxygen during the process of decomposition and, since the water column does not mix, the oxygen in the deeper water is gradually depleted leading, in extreme cases, to deoxygenation. When this happens, most aquatic life becomes confined to the surface waters. Deoxygenation of reservoir waters is particularly common soon after a dam has been constructed, since the filling phase of the reservoir tends to involve inundation of large amounts of terrestrial organic matter. This fuels the development of large populations of microbes that quickly deplete deeper waters of oxygen.

During flood periods, the still waters of the reservoir may be mixed by flows of water from upstream; strong wind blowing across the water surface can have the same effect.

Regardless, such periodic stratification and deoxygenation of deep waters that may occur during much of the year is very different from the continual mixing of surface and bottom waters that occurs in even the deepest rivers. The extent and duration of reservoir stratification is variable and depends on the shape of the reservoir; those constructed in deep valleys are more likely to become stratified and deoxygenated than those that are shallow or have a large surface area relative to their volume.

Downstream effects of dams on rivers can occur immediately downstream of the dam, or may persist for considerable distances. In extreme cases, the dam may hold all water and prevent any flow to escape downstream; this causes dewatering and all aquatic life perishes in the drawdown zone. More commonly, at least a proportion of the natural flow volume is allowed to pass the dam. The condition of this water depends upon whether it has been released from the surface of the reservoir, or if it comes from the reservoir depths. Surface waters are usually clear, warm and oxygen-rich, and contain little detritus (in the form of FPOM) but may have much more phytoplankton than would be found in river waters. Water released from the depths of the reservoir, on the other hand, is cool and oxygen poor, and contains little or no phytoplankton. Accordingly, animals downstream of the dam may have to deal with elevated temperatures but enjoy a rich food supply (in the form of phytoplankton), or may have to cope with cool conditions, low oxygen and little food.

The distance that the effects of a dam are felt downstream depend on the extent to which the temperature of the reservoir waters vary from (above or below) ambient river temperatures, as well as the distance that it takes for filter feeders (such as hydropsychids and simuliids; see Section 6) to consume the phytoplankton. However, there is an additional effect of dams. Regardless of whether water is released from the surface or depths of the reservoir, the water will contain little suspended sediment. Once the water starts to move downhill in the channel below the dam, it has a tendency to erode and transport material from the river bed or banks creating an erosion zone that often extends for several kilometres below the dam. This means that fine sediments, as well as sand and gravel, may be eroded, and these substrates, which may be important to spawning fishes, are often scarce in river sections downstream of dams.

One of the most fundamental biological effects of a dam across a river is the disconnection of migratory pathways by which fish and other animals such as shrimps travel up- and downstream. This may result in the elimination of certain species that make upstream migrations for breeding above the dam, or may cause a drastic decline in their abundance. Dams may also lead to the fragmentation of populations, leading to genetic segregation that may deplete biodiversity (see Section 8). Note that dams do not merely block migrating adult fishes. Species that spawn upstream may rely on the current to transport their eggs and larvae downstream. The slower water in reservoirs means that these fish eggs and larvae that are normally carried by the current in rivers settle to the bottom and may die. Therefore reservoirs and dams pose a barrier to downstream migration of young fish, as well as to the upstream migration of the adults.

Downstream Effects
Downstream effects in rivers can include impacts immediately below the dam, or may persist for considerable distances along the course of the system.

Short Distance Effects. Water discharges into the river downstream of the dam through three sources:

The water used for operating the powerplant in the case of hydroelectric reservoirs, or releases to empty the reservoir after a flood event in the case of flood control reservoirs. (In reservoirs for urban supply and agriculture the water may never return to the river or may do so at a considerable distance downstream.);

Water used to operate any fish-passes associated with the dam; and

Excess water released over the spillway.

The positioning of the spillway is very important in determining the effects of dams on the tailwaters immediately downstream. Spillways situated at the top of the dam discharge highly oxygenated, warm water into the river downstream. Spillways situated near the bottom of the dam release cold, deoxygenated water.


Dam spillwayPhysical, chemical and biological changes occur in the water stored in reservoirs. As a result the water discharged at a dam can be of different chemical composition and show a different seasonal pattern to that of the natural river. Generally, the reservoir acts as a nutrient sink, storing much of the inflowing nitrogen and phosphorus in the sediment. Some chemicals may increase, for example, in arid regions the water may become saltier through evaporation, particularly in areas where the inflowing streams have a high salt content.

Reservoirs also influence the temperature of the river immediately below the dam and, although this alters the ambient temperature as the water moves downstream, the thermal impact of the dam can be detected for a considerable distance downstream. Water temperature is important for the assessment of impacts of a dam and reservoir on downstream aquatic habitats, because it influences many important physical, chemical and biological processes. As a result, the thermal change may have significant effects on the living organisms in the river.

The quality of water discharged from a stratified reservoir depends on the position of the outflow relative to the different water layers in the reservoir. When the water is stratified, water released from near the surface will be well oxygenated, warm and depleted in nutrients. Water released from near the bottom will be cold, oxygen-depleted and nutrient-rich. It may also contain toxic contaminants such as hydrogen sulphide, iron and manganese.

In both upper and lower intakes, the water is likely to have a low silt loading, although water discharged from near the bottom is more likely to contain some silt. The low silt loading relative to flow means that the water has a tendency to pick up more suspended matter from the bottom of the river, leading to an erosion zone that often extends for a considerable distance below the dam. The bottom of the river is cut down rapidly in this erosion zone, and often bottom substrates such as gravel, important for the spawning of fish, are carried away.

Long Distance Effects. The most persistent effect of dams is the alteration to the natural flow regimes that is described in Section 4. The flood peak is suppressed, and dry season flows increased to even out the hydrograph downstream. These impacts can extend throughout the length of the river in the case of a single, mainstream dam. However, the effects of smaller dams on tributary flows may be local only in the case where flow in the main river may be sustained by natural flow regimes from other tributaries. Problems arise when all tributaries are dammed, as the cumulative effects of many small dams may be as severe as the effects of a major mainstream dam. Similarly, cascades of dams may completely alter the flow characteristics of the river on which they are built.


The changes downstream of dams may include (but are not confined to) the following:

• Alteration of the timing, duration and magnitude of flood peaks and low-flow periods;

• Modification of river temperature that may influence the timing or initiation of reproduction and breeding migrations by aquatic animals;

• Declines in the amounts of FPOM and suspended inorganic material and reductions in the distances over which they are transported;

• Reductions in the extent and duration of floodplain inundation, and modification of the flood-pulse resulting in reductions in land-water exchanges of material.

The combined effects of dams can depend upon where they are constructed along the course of a river, and have been described in the discussion of the Serial Discontinuity Concept (see Section 7). Increasing awareness of the detrimental effects of dams on river ecology has focussed attention on the need to release water from dams in a way that mimics the natural pattern of flood peaks and low flow periods, as well as allocation of sufficient water to maintain the ecological function of areas downstream. The latter has given rise to attempts to estimate and maintain the minimum flows needed to achieve this goal. Even where minimum flows can be estimated accurately (and this is certainly something that is difficult to calculate), allocation of sufficient water to ecological functions is a major challenge for river managers since water allocated to one function (e.g. irrigation) is usually unavailable for another (e.g. maintaining river health). Furthermore the economic benefits of increased irrigation, of flood control, or of hydroelectric power generation are relatively easy to measure, whereas the benefits of maintaining ecological function are often less tangible.


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